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While Elton and his peers observed that less diverse communities are characterized by greater fluctuations in population density, the theory remained largely untested.
In 1973, Robert May published a mathematically-based study that seemed to contradict the long-held hypothesis. May reported that communities with higher diversity tended to be less, not more, stable because higher diversity tended to undermine individual species. By the 1990s, field studies by Tilman et al. raised the possibility that the number of species in an ecosystem might greatly influence how that ecosystem functions by showing that the stabilizing effect of diversity on the whole ecosystem was much greater than the destabilizing effect of diversity on an individual species...
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