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These will be examined in order.
When an actor's behaviour increases the fitness of the behaviour's receiver at the expense of the actor, then the behaviour is said to be altruistic. This definition is simplistic but provides a good starting point; throughout the discussion it will be sharpened. In contrast to so-called selfish behaviour increases the actor's fitness while reducing the fitness of the receiver. If both the actor and the receiver gain in fitness following a behaviour, then the behaviour is cooperative. We must keep in mind that these costs and benefits are measures of reproductive success, the number of offspring surviving to reproductive age and thus the number of genes passed to the next generation...
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