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He says that learning should be studied in the framework of representation. Studying animals, he developed a framework that linked quantities computed and stored by the nervous system representing aspects of the animal's environment to the animal's relationship with that environment. The brain is said to represent an aspect of the environment when there is a functioning isomorphism between an aspect of the environment and a brain process that adapts the animal's behavior to it.
Again he uses the mathematical definition of isomorphism saying that they are formal correspondences between distinct systems of mathematical study; often with one being an object not well understood and another more richly developed system...
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